Peer-reviewed papers


Cadena-Castañeda O, Landeck I, Hemp C (2024) Studies on chevron crickets: Minarisoma gen. nov. (Orthoptera: Anostostomatidae), a new genus and a new species from South Africa. Journal of Orthoptera Research 33(1):147–156.

Heller K-G, Hemp C (2024): Egg shape and size in Phaneropterinae and other Tettigonioidea (Orthoptera: Ensifera): A global review with new data. Journal of Orthoptera Research 33(1): 103-112.

Hemp A, Hemp J (2024) Weather or not—Global climate databases: Reliable on tropical mountains? PLoS ONE 19(3): e0299363.

Bianco G, Manning P, Schleuning M (2024) A quantitative framework for identifying the role of individual species in Nature’s Contributions to People. Ecology Letters, 27, e14371.

Cadena-Castañeda O, Hemp C (2024) Studies on chevron crickets: Tryposoma gen. nov. (Orthoptera: Anostostomatidae), a new genus from Tanzania. Journal of Orthoptera Research 33(1): 59–66.

Pearson J, Gross M, Hofmann J (2024) Non-material contributions of nature expressed by former tourists of Mount Kilimanjaro, Tanzania. People and Nature (online early).

Publications from previous years


Ma H, Crowther TW, Mo L, […], Fischer M, […], Hemp A, […], Zohner CM (2023) The global biogeography of tree leaf form and habit. Nature Plants 9: 1795-1809.

Darbyshire I, Hemp A (2023) A further new species of Isoglossa (Acanthaceae) from the eastern Arc Mountains of Tanzania. Kew Bulletin 78: 499-507.

Delavaux CS, Crowther TW, Zohner CM, […], Fischer M, […], Hemp A, […], Maynard DS (2023) Nature diversity buffers against severity of non-native tree invasions. Nature 621: 773-781. 

Gross M, Pearson J, Arbieu U, Riechers M, Thomsen S, Martín-López B (2023) Tourists’ valuation of nature in protected areas: a systematic review. Ambio 52: 1065-1084.

Hemp A, Bianco G, Pócs T (2023) Homegardens and coffee plantations on Kilimanjaro: Refugium for bryophytes in densely cultivated landscapes. Hattoria 14: 1-16.

Hemp C (2023) Acrida bara, synomymous with A. sulphuripennis (Orthoptera, Acrididae, Acridinae). Journal of Orthoptera Research 32: 115-117.

Hemp C (2023) A new species of Neoleva (Caelifera, Acridoidea, Acrididae, Gomphocerinae) from Central Tanzania. Journal of Orthoptera Research 32: 127-131.

Hemp C, Montealegre-Z F, Woodrow C, Heller K-G (2023) Bush-crickets with very special ears and songs – review of the East African Phaneropterinae genus Dioncomena Brunner von Wattenwyl, 1878, with notes on its biogeography and the description of new species. Deutsche Entomologische Zeitschrift 70: 221-259.

Kaasalainen U, Kirika PM, Mollel NP, Hemp A, Rikkinen J (2023) The lichen genus Sticta (Lobariaceae, Peltigerales) in East African montane ecosystems. Journal of Fungi 9: 246.

Watts M, Mpanda M, Hemp A, Peh K (2023) The potential impact of future climate change on the production of a major food and cash crop in tropical (sub)montane homegardens. Science of the Total Environment 865: 161263.


Heller K-G, Chobanov D, Warchalowska-Sliwa E, Hemp C (2022) Review of song patterns and sound production in armoured ground crickets (Orthoptera: Tettigoniidae: Hetrodini) with karyological data and taxonomic notes. Zootaxa 5120: 451–481.                          

Hemp A, Philipp C, Hemp C (2022) European Union’s Natura 2000 network: an effective tool for nature conservation? The relic pine forests of the Franconian Jura. Biodiversity and Conservation 31: 1909–1926.

Hemp C, Rowell H F (2022) A new species of Burttia Dirsh, 1951 (Caelifera, Acridoidea, Acrididae, Catantopinae) from the Eastern Arc Mountains of Tanzania. Journal of Orthoptera Research 31: 1–7.

Kaasalainen U , Biermann L, Mollel NP, Schmidt AR, Hemp A (2022) Peltigera (Lecanoromycetes) on Mt Kilimanjaro, East Africa. The Lichenologist 54: 231–243.

Komposch A, Ensslin A, Fischer M, Hemp A (2022) Aboveground deadwood biomass and composition along elevation and land-use gradients at Mount Kilimanjaro. Frontiers in Ecology and Evolution 9: 732092.

París M, Hemp C (2022) The type of Macroscirtus bicolor Bolívar, 1893 (Orthoptera, Ensifera, Tettigoniidae, Mecopodinae). Graellsia 78: e170.

Vogeler A-VB, Otte I, Ferger S, Helbig-Bonitz M, Hemp A, Nauss T, Böhning-Gaese K, Schleuning M, Tschapka M, Albrecht J (2022) Associations of bird and bat species richness with temperature and remote sensing-based vegetation structure on a tropical mountain. Biotropica 54: 135-145.

Ziegler A, Meyer H, Otte I, Peters M K, Appelhans T, Behler C, Böhning-Gaese K, Classen A, Detsch F, Deckert J, Eardley C D, Ensslin A, Ferger S W,  Fischer M, Frederiksen S B, Gebert F, Haas M, Helbig-Bonitz M, Hemp A, Hemp C, Kakengi V, Kalko E K V,  Kindeketa W J, Kleyer M, Mayr A, Mwangomo E,  Ngereza C, Renner M, Reudenbach C, Röder J, Rutten G, Schellenberger Costa D, Schleuning M, Stefan-Dewenter I, Tschapka M, Vollstädt M G R, Peters R S, Ssymank A, Tardanico J, Wöllauer S, Zancolliy G, Zhang J, Brandl R, Nauss T (2022) Potential of Airborne LiDAR Derived Vegetation Structure for the Prediction of Animal Species Richness at Mount Kilimanjaro. Environmental Remote Sensing 14: 786. 


Byamungu RM, Schleuning M, Ferger SW, Helbig-Bonitz M, Hemp A, Neu A, Vogeler A, Böhning-Gaese K, Tschapka M, Albrecht J (2021) Abiotic and biotic drivers of functional diversity and functional composition of bird and bat assemblages along a tropical elevation gradient. Diversity and Distributions 27: 2344-2356.

Cuni-Sanchez A, Sullivan MJP, Platts PJ, […], Hemp A, […], Zibera E (2021) High aboveground carbon stock of African tropical montane forests. Nature 596: 536-542.

Grzywacz B, Warchałowska‑Śliwa E, Kociński M, Heller K-G, Hemp C (2021) Diversification of the Balloon bushcrickets (Orthoptera, Hexacentrinae, Aerotegmina) in the East African mountains. Scientific Reports 11: 9878.

Heller K-G, Heller M, Samietz J, Volleth M, Hemp C (2021) Similar songs, but divers mate localization strategies in the three Phaneroptera species occurring in Western Europe (Orthoptera: Tettigonioidea). European Journal of Entomology 118: 111-122.

Hemp A, Del Fabbro C, Fischer M (2021) Area modulates the effect of elevation but not of land use or canopy on tropical plant species richness. Biodiversity and Conservation 30: 4265-4277.

Hemp C (2021) A new species of Physocrobylus (Caelifera, Acridoidea, Acrididae, Coptacrinae), with notes on the phenology and habitat of the genus. Journal of Orthoptera Research 30: 1-6.

Hemp C, Massa B (2021) Biogeographical and evolutionary aspects of a Guineo-Congolian bushcricket tribe: revision of the genera Cestromoecha Karsch, 1893 and Poreuomena Brunner von Wattenwyl, 1878, with the description of new species (Orthoptera, Tettigoniidae, Phaneropterinae). Deutsche Entomologische Zeitschrift 68: 45-79.

Hemp C, París M (2021) A new species of Apteroscirtus Karsch, 1891 (Ensifera, Tettigoniidae, Mecopodinae) from Angola. Zootaxa 5052: 287-291.

Kaasalainen U, Tuovinen V, Kirika PM, Mollel NP, Hemp A, Rikkinen J (2021) Diversity of Leptogium (Collemataceae, Ascomycota) in East African montane ecosystems. Microorganisms 9: 314.

Mollel NP, Hemp A, Fischer M (2021) Phylogenetic diversity of plant communities in relation to elevation and human impact at Mount Kilimanjaro, Tanzania. Tanzania Journal of Forestry and Nature Conservation 90: 54-66.

Prunera-Olivé J, Vorontsova MS, Williams EV, Mollel NP, Hemp A (2021) Checklist of Kilimanjaro grasses shows that both plot and herbarium methods are necessary to record diversity. Phytotaxa 501: 201-244.

Seibold S, Rammer W, Hothorn, T, […], Hemp A, Hemp C, […], Müller J (2021) The contribution of insects to global forest deadwood decomposition. Nature 597: 77-81.

van der Plas GW, Rucina SM, Hemp A, Marchant RA, Hooghiemstra H, Schüler L, Verschuren D (2021) Climate-human-landscape interaction in the eastern foothills of Mt. Kilimanjaro (equatorial East Africa) during the last two millennia. The Holocene 31: 556-569.

Warchałowska-Śliwa E, Grzywacz B, Kociński M, Maryańska-Nadachowska A, Heller K-G, Hemp C (2021) Highly divergent karyotypes and barcoding of the East African genus Gonatoxia Karsch (Orthoptera: Phaneropterinae). Scientific Reports 11: 22781.


Bodé S, de Wispelaere L, Hemp A, Verschuren D, Boeckx P (2020) Water‐isotope ecohydrology of Mount Kilimanjaro. Ecohydrology 13: e2171.

Classen A, Eardley CD, Hemp A, Peters MK, Peters RS, Ssymank A, Steffan-Dewenter I (2020) Specialization of plant-pollinator interactions increases with temperature at Mt. Kilimanjaro. Ecology and Evolution 10: 2182-2195.

Heller K-G, Hemp C (2020) Hyperdiverse songs, duetting, and the roles of intra- and intersexual selection in the acoustic communication of the genus Eurycorypha (Orthoptera: Tettigonioidea, Phaneropterinae). Organims Diversity & Evolution 20: 597-617.

Hemp C (2020) A new species of Stenampyx Karsch, 1890 from East Africa (Orthoptera: Tettigoniidae, Pseudophyllinae, Phyllomimini) – evidence of a former connection of West-Central and East African forests. Zootaxa 4763: 593-599.

Hemp C (2020) New species of Mecopodinae (Orthoptera, Tettigoniidae) from Tanzania. Zootaxa 4790: 138-150.

Hemp C, Scherer C, Brandl R, Pinkert S (2020) The origin of the endemic African grasshopper family Lentulidae (Orthoptera: Acridoidea) and its climate-induced diversification. Journal of Biogeography 47: 1805-1815. https://doi. org/10.1111/jbi.13880

Molina-Venegas R, Fischer M, Hemp A (2020) Plant evolutionary assembly along elevational belts at Mt. Kilimanjaro: using phylogenetics to assess biodiversity threats under climate change. Environmental and Experimental Botany 170: 103853.

Molina-Venegas R, Fischer M, Mollel NP, Hemp A (2020) Connecting plant evolutionary history and human well-being at Mt. Kilimanjaro, Tanzania. Botanical Journal of the Linnean Society 194: 397-409.

Ndossi EM, Becker JN, Hemp A, Dippold MA, Kuzyakov Y, Razavi BS (2020) Effects of land use and elevation on the functional characteristics of soil enzymes at Mt. Kilimanjaro. European Journal of Soil Biology 97: 103167.

Shen C, Gunina A, Luo Y, Wang J, He J-Z, Kuzyakov Y, Hemp A, Classen AT, Ge Y (2020) Contrasting patterns and drivers of soil bacterial and fungal diversity across a mountain gradient. Environmental Microbiology 22: 3287-3301.

Sierra Cornejo N, Hertel D, Becker JN, Hemp A, Leuschner C (2020) Biomass, morphology, and dynamics of the fine root system across a 3,000-m elevation gradient on Mt. Kilimanjaro. Frontiers in Plant Science 11: 13.

Vollstädt MGR, Albrecht J, Böhning-Gaese K, Hemp A, Howel KM, Kettering L, Neu A, Neuschulz EL, Quitián M, Santillán VE, Töpfer T, Schleuning M, Fritz SA (2020) Direct and plant-mediated effects of climate on bird diversity in tropical mountains. Ecology and Evolution 10: 14196-14208.

Warchałowska-Śliwa E, Grzywacz B, Maryańska-Nadachowska A, Heller K-G, Hemp C (2020) Rapid chromosomal evolution in the bush-cricket Gonatoxia helleri (Orthoptera: Phaneropterinae). Comparative Cytogenetics 14: 417-435.